Do small tributaries function as refuges from floods? A test in a salmonid-dominated mountainous river

Excerpts from the text: On 8–10 August 2003, a powerful typhoon hit Hokkaido Island, Japan, accompanied with heavy rain, which allowed us to investigate the potential role of tributaries as refuges from flooding. We had just completed annual population census in four small tributaries of a river system 1–2 days before the typhoon.... Overall, our results did not support the hypothesis that many large fishes immigrate to small tributaries during floods. ... Despite the lack of evidence of mass movement, our result suggested a few immigrants from the main stem (i.e., juvenile white-spotted charr, sculpin and a few relatively large Dolly Varden). Because more than 100 small tributaries exist in the Shiisorapuchi River (Koizumi 2011), only a few individuals escaping to each tributary should accumulate to a great number enough to re-colonise main stem habitats even if fishes in the main stem were extirpated. Multiple refuges at different spatial scales should increase resistance and ⁄ or resilience of fish populations (Sedell et al. 1990; Pearsons et al. 1992). Thus, the roles of tributaries as refuges would deserve further attention